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Trans-ethnic association study of blood pressure determinants in over 750,000 individuals.

Nat Genet · 2018
L1 No data access 2/4
Why this verdict

Part of the results reproduced; minor but material deviations remained.

Reproduced on the brainbox compute brainarbeit.com
Scoring basis — itemised

Every item that counted toward this verdict, and the exact part of the reproduction that produced it.

Supporting (toward a concern)
Content-critical question only partially held
+2 pts
From: Q5 · Derivability / plausibility 🟡
Content-critical question only partially held
+2 pts
From: Q7 · Core claim 🟡
Content-critical question only partially held
+2 pts
From: Q8 · Severity of the miss (overall human judgment) 🟡
Minor / cosmetic deviation
+1 pts
From: Q3 · Location of the main deviation 🟡
Minor / cosmetic deviation
+1 pts
From: Q6 · Severity of the deviation 🟡
Input / endpoint not comparable 1:1
+1 pts
From: Q1 · Data identity 🔴
Concordant (toward reproduced)
Code + data deposited & functional
-2 pts
From: Data & code availability Available & functional
Total score +7
✓ What held up
  • No authors-side cause for any deviation
What did not (or only partly)
  • 🔴Could not use the authors’ exact input data
  • 🔴Reported values were only indirectly comparable
  • 🟡A deviation arose in the data or preprocessing
  • 🟡Reported values were not (fully) derivable from the shared data
  • 🟡The deviation was non-trivial in magnitude
  • 🟡The central claim did not (fully) hold under reproduction
  • 🟡Overall, the reproduction showed a material discrepancy
No data access Data access not granted

This paper has a computational component, but its primary data is legally or ethically access-restricted — identifiable patient cohorts, rare-disease genomes, or controlled-access biobanks that cannot be openly shared. The reproduction therefore could not be attempted. That is a neutral verdict: it does not mean the result is wrong or that the authors fell short — only that, for legitimate privacy reasons, it cannot be independently checked from public data. We deliberately do NOT assign a 0–100 score here, because a low number would wrongly read as a failed reproduction.

Reproduction agent’s raw note

DROP (data_restricted). Paper: 'Trans-ethnic association study of blood pressure determinants in over 750,000 individuals' (PMID 30578418, doi 10.1038/s41588-018-0303-9) is a multi-cohort trans-ethnic GWAS meta-analysis of systolic/diastolic/pulse blood pressure. The pipeline-derived results (per-variant association statistics, trans-ethnic meta-analysis, novel-locus discovery) require INDIVIDUAL-LEVEL genotype + phenotype data from controlled/application-only cohorts — primarily UK Biobank (application-only, MTA) and the Million Veteran Program (dbGaP controlled-access) plus consortium cohorts. That input is not publicly obtainable, so the end-to-end association pipeline cannot be re-run for reproduction. The link-mined code artifact (github.com/gabraham/flashpca) is only a sub-component (fast PCA for ancestry/population-structure covariates), not the GWAS/meta-analysis pipeline, and the link-mined data accession (GEO GSE107585, a kidney single-nucleus RNA-seq dataset) is a text-mining/enrichment false positive — it is not the GWAS input and cannot pin any reported BP-association value. NOT ATTEMPTED: no «our HPC» compute submitted, because no public input data exists to feed a faithful reproduction; fabricating a result to avoid the drop is explicitly disallowed by the brief. Published GWAS summary statistics may exist downstream, but re-deriving them is the restricted-data step. Verdict provisional for human audit.

These records describe the outcome of reproduction attempts carried out autonomously by brainbox using large language models (LLMs). They are not peer review, not an audit, and not a determination of error or misconduct by any author. A verdict reflects what one attempt could or could not reproduce — which may depend on data access, undocumented parameters, the computing environment, or the depth of effort — and not a judgement of the people who did the work. We can be wrong, and we correct mistakes quickly: every record carries a “report an error” button.

Assessment versions

Every reproduction run is kept as an immutable version — anchored to the data as it stood, with a tamper-evident chain hash. A rerun (e.g. after an author updates a deposit) adds a new version; the previous one stays on record.

  1. v1 current initial assessment
    assessed: 2026-06-16 ⛓ 8cb8426af00b
✎ I am an author of this paper

Updated or fixed a deposit, or is there an erratum? Ask us to re-run the metrics. We verify by email first; the new result is published as a new version with full history — nothing is overwritten.

Reason for the rerun

We email you a confirmation link first. The rerun is an objective re-measurement — it cannot change the verdict in your favour, only ask us to look again.

Provenance — full disclosure

When this reproduction was carried out, which methodology version was used, and by whom — so the record can be audited and checked independently.

Reproduced
2026-06-16
Rubric version
v1.0
Assessed by
🤖 AI curator · claude (ai-curator room) · v1.0 · run #1 2026-06-16
no human curator yet
Last updated
2026-08-05

Provisional, curator- or AI-assessed, and independently checkable. A reproduction outcome states what one attempt could reproduce — not a judgement of the authors.

Deep full-text extraction

Model: opus
Founding hypothesis

Can a trans-ethnic, multi-omic analysis of blood pressure in over 750,000 individuals reinterpret the genetic architecture of blood pressure to identify novel genes, tissues, phenome, and medication contexts underlying blood pressure homeostasis?

Core claims
  • Discovery and replication GWAS of SBP, DBP and pulse pressure in up to 776,078 individuals identified 208 novel common blood pressure SNPs and 53 rare variants. finding
  • A transcriptome-wide association study detected 4,043 blood pressure associations with genetically-predicted expression of 840 genes across 45 tissues. finding
  • Murine renal single-cell RNA sequencing identified upregulated blood pressure genes in kidney tubule cells. finding
  • 505 independent loci (201 novel, 304 previously reported) were associated with one or more blood pressure traits. finding
  • Ten rare missense variants from seven genes (PDE3A, SLC9A3R2, RRAS, PHC3, DBH, COL21A1, NOX4) were associated with blood pressure traits. finding
  • Trans-ethnic meta-analysis combined with TWAS, PheWAS of genetic risk scores, pathway/tissue enrichment and gene-drug analyses constitutes the multi-omic framework for blood pressure homeostasis. method
  • Effect-size correlations between white, black, and Hispanic groups in MVP were weaker than previously reported, though directions of effect were largely consistent. finding
  • Most blood pressure SNPs are noncoding and reside in regulatory elements, motivating gene-based imputed-expression association tests. mechanism
Experimental setups
Assay System Perturbation Readout Platform
GWAS common variant meta-analysis (single variant, MAF >1%) Human; MVP + UK Biobank discovery, ICBP + BioVU replication none SBP, DBP and pulse pressure association (mmHg per allele)
GWAS rare exonic variant analysis (MAF <1%) Human; MVP discovery, BioVU + BP-ICE exome chip replication none blood pressure trait association of missense variants exome chip
Genome-wide rare variant analysis Human; MVP discovery, UK Biobank replication none rare variant blood pressure association
Transcriptome-wide association study (TWAS / genetically-predicted gene expression) Human; 45 tissues none association of genetically-predicted gene expression with blood pressure (840 genes)
Phenome-wide association study (PheWAS) Human; MVP blood pressure genetic risk scores clinical phenome associations
Single-cell RNA sequencing Mouse (murine) kidney none cell-type-specific expression of blood pressure genes (kidney tubule cells)
Pathway and tissue gene set enrichment analysis Human none enriched pathways and tissues
Conditional analysis of rare variants on sentinel common variants Human; MVP whites discovery sample none effect-size change after conditioning
Key results
  • 208 novel common blood pressure SNPs and 53 rare variants discovered
  • 4,043 blood pressure associations with genetically-predicted expression of 840 genes across 45 tissues
  • Blood pressure genes upregulated in murine kidney tubule cells
  • 505 independent loci identified (201 novel, 304 previously reported); replicated novel loci included 124 SBP, 4 DBP, 123 pulse pressure
  • Novel loci had smaller per-allele effects (0.24, 0.14, 0.18 mmHg for SBP, DBP, PP) than known loci (0.32, 0.27, 0.27 mmHg) 0.24/0.14/0.18 vs 0.32/0.27/0.27 mmHg per allele
  • Sentinel SNPs at all loci explained 3.56%, 1.06%, 3.72% of variance for SBP, DBP, pulse pressure; novel variants contributed 0.80%, 0.24%, 0.72% 3.56%/1.06%/3.72%
  • Ten missense variants from seven genes associated with blood pressure; mean absolute effects 1.52, 0.63, 1.50 mmHg per allele for SBP, DBP, PP 1.52/0.63/1.50 mmHg per allele
  • rs139491786 (SLC9A3R2) showed >50% effect reduction after conditioning on common variant rs140869992 >50% reduction; r2=0.35
Key statistics
  • count 776,078 participants (up to) (total GWAS sample for SBP, DBP, pulse pressure)
  • count 459,777 meta-analyzed in discovery (318,891 MVP + 140,886 UKB) (discovery sample)
  • count 316,301 replication participants (ICBP + BioVU) (common variant replication)
  • count 445,360 UKB replication for genome-wide rare variants; BP-ICE Nmax 420,704 (rare variant replication)
  • pvalue meta-analysis P < 5 × 10−8 (genome-wide significance threshold)
  • fold_change 0.32, 0.27, 0.27 mmHg per allele (known loci) (mean effect of trait-increasing alleles SBP/DBP/PP)
  • other r2 = 0.35 (LD between rs139491786 and rs140869992 in SLC9A3R2)
  • other 13% of annual deaths worldwide attributable to elevated blood pressure (background burden estimate)

Statistical methods review

Model: opus

A neutral, descriptive read of the statistical approach — what was done, and (for shared learning, not as criticism) what could also have been done.

This is a large trans-ethnic genome-wide association meta-analysis of systolic, diastolic and pulse pressure in up to ~776,078 participants, using a discovery (MVP + UK Biobank, N=459,777) and independent replication (ICBP, BioVU, BP-ICE) design for common and rare variants. Single-variant association testing was combined via fixed-effect-style meta-analysis, with locus novelty and replication defined by pre-specified P-value, distance, and linkage-disequilibrium thresholds; conditional analyses identified independent secondary signals, and downstream transcriptome-wide association, PheWAS, pathway/tissue enrichment, and single-cell expression analyses were performed. Results were reported primarily as per-allele effect estimates (mmHg) and P-values, with variance-explained summaries for the identified loci.

Replicationunclear Sample sizeSample sizes stated by cohort/stage (e.g., 318,891 MVP, 140,886 UKB, 316,301 ICBP/BioVU replication, up to 776,078 total); no formal power calculation described in the provided text GroupsGenotype-blood pressure associations (SBP, DBP, pulse pressure) within and across ancestry groups; discovery vs replication cohorts Pairingna Randomization/blindingna DispersionSD Exact p-valuesno Effect sizesyes Multiplicity correctionGenome-wide significance threshold (P < 5 × 10⁻⁸) plus staged replication criteria (discovery P < 1 × 10⁻⁶, replication P < 0.05, consistent direction, combined P < 5 × 10⁻⁸)
Statistical tests used
Test Applied to n Assumptions
Single-variant genome-wide association test (per-trait, genome-wide significance threshold P < 5 × 10⁻⁸) Discovery and meta-analysis of common variants for SBP, DBP and pulse pressure up to 459,777 discovery; up to 776,078 combined as stated not stated
Meta-analysis combining discovery and replication association statistics Common-variant replication (ICBP/BioVU) and rare-variant replication (UKB, BP-ICE) 316,301 common-variant replication; up to 420,704 (BP-ICE) and 445,360 (UKB) rare-variant replication not stated
Conditional (single-variant conditional) association analysis Identification of conditionally independent secondary signals and conditioning rare exonic variants on sentinel common variants in MVP whites 318,891 (MVP discovery) as stated not stated
Correlation of per-allele effect estimates across ancestry groups Trans-ancestry comparison of known and novel loci across white, black, and Hispanic MVP samples not stated
Transcriptome-wide / gene-based association of genetically predicted gene expression with blood pressure TWAS detecting 4,043 associations for 840 genes across 45 tissues not stated
Phenome-wide association study (PheWAS) of blood pressure genetic risk scores; pathway and tissue gene-set enrichment analyses Blood pressure clinical phenome in MVP and downstream enrichment analyses not stated
Approaches that could also have been used
  • Replication and significance were defined using fixed P-value thresholds (e.g., discovery P < 1 × 10⁻⁶, replication P < 0.05, combined P < 5 × 10⁻⁸).
    Could also: Reporting effect estimates with 95% confidence intervals alongside the threshold-based decisions. — Confidence intervals convey the precision and plausible range of each per-allele effect, which complements threshold-based significance and aids cross-study comparison.
  • Cross-ancestry consistency was assessed by correlating per-allele effect estimates across white, black, and Hispanic groups.
    Could also: A formal trans-ethnic meta-analysis framework (e.g., MANTRA or a random-effects/heterogeneity model such as Cochran's Q or I²). — Such methods explicitly model and quantify between-ancestry effect heterogeneity, adding a formal test statistic to accompany the descriptive effect-size correlations.
  • Variant-level multiplicity was handled via the conventional genome-wide threshold of 5 × 10⁻⁸.
    Could also: Reporting a study-specific false discovery rate (e.g., Benjamini-Hochberg) for the families of downstream tests (TWAS, PheWAS, enrichment). — An FDR summary across the many downstream comparisons would provide an additional, explicitly stated control on the expected proportion of false positives within each analysis family.
  • Independent secondary signals were identified using single-variant conditional analysis.
    Could also: Approximate conditional and joint analysis (e.g., GCTA-COJO) or fine-mapping approaches that yield credible sets. — These approaches can jointly model multiple signals and provide posterior probabilities or credible intervals for likely causal variants, adding resolution to the independent-signal characterization.
  • Descriptive cohort characteristics were summarized as mean (standard deviation).
    Could also: Also presenting medians with interquartile ranges for skewed measures. — Median/IQR can convey the distribution shape for variables that may be non-normal, complementing the mean and SD already reported.

Result convergence & founder nodes

Findings this paper shares with others that ran a comparable experiment. A node’s strength is how many independent papers report it (replication breadth) — not how often it is cited, so a heavily-replicated but under-cited founder still stands out.

Citation network

Where this publication sits in the reproducibility-weighted citation graph — what it is built on, and what is built on it. Citation data from OpenAlex.

Authors · 100
1Understanding Society Scientific Group 2Ayush Giri 3Million Veteran Program 4Jacklyn N. Hellwege 5Jacob M. Keaton 6Jihwan Park 7Chengxiang Qiu 8Helen Warren 9Eric S. Torstenson 10Csaba P. Kövesdy 11Yan V. Sun 12Otis D. Wilson 13Cassianne Robinson‐Cohen 14Christianne L. Roumie 15Cecilia P. Chung 16Kelly A. Birdwell 17Scott M. Damrauer 18Scott L. DuVall 19Derek Klarin 20Kelly Cho 21Yu Wang 22Εvangelos Εvangelou 23Claudia P. Cabrera 24Louise V. Wain 25Rojesh Shrestha 26Brian S. Mautz 27Elvis A. Akwo 28Muralidharan Sargurupremraj 29Stéphanie Debette 30Michael Boehnke 31Laura J. Scott 32Jian’an Luan 33Jing-Hua Zhao 34Sara M. Willems 35Sébastien Thériault 36Nabi Shah 37Christopher Oldmeadow 38Peter Almgren 39Ruifang Li‐Gao 40Niek Verweij 41Thibaud Boutin 42Massimo Mangino 43Ioanna Ntalla 44Elena V. Feofanova 45Praveen Surendran 46James P. Cook 47Savita Karthikeyan 48Najim Lahrouchi 49Chunyu Liu 50Nuno Sepúlveda 51Tom G. Richardson 52Aldi T. Kraja 53Philippe Amouyel 54Martin Farrall 55Neil R Poulter 56Markku Laakso 57Eleftheria Zeggini 58Peter Sever 59Robert A. Scott 60Claudia Langenberg 61Nicholas J. Wareham 62David Conen 63Colin Neil Alexander Palmer 64John Attia 65Daniel I. Chasman 66Paul M. Ridker 67Olle Melander 68Dennis Owen Mook-Kanamori 69Pim van der Harst 70Francesco Cucca 71David Schlessinger 72Caroline Hayward 73Tim D. Spector 74Marjo-Riitta Jarvelin 75Branwen J. Hennig 76Nicholas J. Timpson 77Wei-Qi Wei 78Joshua Smith 79Yaomin Xu 80Michael E. Matheny 81Edward D. Siew 82Cecilia M. Lindgren 83Karl‐Heinz Herzig 84George Dedoussis 85Joshua C. Denny 86Bruce M. Psaty 87Joanna M. M. Howson 88Patricia B. Munroe 89Christopher Newton‐Cheh 90Mark J. Caulfield 91Paul Elliott 92J. Michael Gaziano 93John Concato 94Peter W.F. Wilson 95Philip S. Tsao 96Digna R. Velez Edwards 97Katalin Suszták 98Christopher J. O’Donnell 99Adriana M. Hung 100Todd L. Edwards
Citations
500
Impact: very high
Foundation confidence
None of its references are in our reproducibility record yet — its foundation cannot be assessed.
Topics

No assessed neighbours yet — the network grows as more papers are assessed.

Data lineage

The datasets this paper uses (text-mined from the full text via Europe PMC), and which other assessed papers stand on the same data. A shared dataset is a factual link — not a judgement.

112410 OMIM in Discussion (http://purl.org/orb/Discussion)
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AF086203 ENA in Table (http://semanticscience.org/resource/SIO_000419)
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CHEMBL355497 ChEMBL in Table (http://semanticscience.org/resource/SIO_000419)
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GSE107585 GEO in Supplementary material (http://purl.obolibrary.org/obo/IAO_0000326)
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phs001672 dbGaP in Supplementary material (http://purl.obolibrary.org/obo/IAO_0000326)
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rs10260816 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs1061808 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs11039216 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs11105354 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs115079907 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs11617448 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs11783703 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs12035750 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs12203179 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs12449170 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs12656497 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs12705390 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs12952051 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs138582164 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs139341533 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs139491786 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs140473396 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs140869992 RefSNP in Results (http://purl.org/orb/Results)
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rs141325069 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs141328069 RefSNP in Results (http://purl.org/orb/Results)
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rs1800629 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs200999181 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs202102042 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs2071382 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs216172 RefSNP in Discussion (http://purl.org/orb/Discussion)
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rs2240716 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs2643826 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs2764043 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs3025380 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs3085380 RefSNP in Results (http://purl.org/orb/Results)
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rs34868542 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs35979968 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs374292503 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs3796592 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs4656180 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs573455 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs58068637 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs6026739 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs60691990 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs6090040 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs61760904 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs6503413 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs6595838 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs6669371 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs73181210 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs74181299 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs80335285 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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rs9603376 RefSNP in Table (http://semanticscience.org/resource/SIO_000419)
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What was reproduced

The exact results taken into scope, with each reported value next to the value our attempt produced.

No individual results have been recorded for this entry yet.

Assessments & scoring basis

Each contributor’s verdict, the per-question basis, and the auditable, itemised worksheet behind it.

🤖 AI curator · claude (ai-curator room) · v1.0 L1 44/100

An automated assessment. It can flag an open question for review but can never, on its own, record a discrepancy verdict (C5) against a paper.

🔴1. Data identity
🔴2. Endpoint comparability
🟡3. Location of the main deviation
🟢4. Cause of the deviation
🟡5. Derivability / plausibility
🟡6. Severity of the deviation
🟡7. Core claim
🟡8. Severity of the miss (overall human judgment)
Scoring basis — itemised

Every item that counted toward this verdict, and the exact part of the reproduction that produced it.

Supporting (toward a concern)
Content-critical question only partially held
+2 pts
From: Q5 · Derivability / plausibility 🟡
Content-critical question only partially held
+2 pts
From: Q7 · Core claim 🟡
Content-critical question only partially held
+2 pts
From: Q8 · Severity of the miss (overall human judgment) 🟡
Minor / cosmetic deviation
+1 pts
From: Q3 · Location of the main deviation 🟡
Minor / cosmetic deviation
+1 pts
From: Q6 · Severity of the deviation 🟡
Input / endpoint not comparable 1:1
+1 pts
From: Q1 · Data identity 🔴
Concordant (toward reproduced)
Code + data deposited & functional
-2 pts
From: Data & code availability Available & functional
Total score +7

This is a data_restricted drop: the trans-ethnic BP GWAS meta-analysis (PMID 30578418) depends on individual-level genotype/phenotype data from controlled-access cohorts (UK Biobank, Million Veteran Program) that are not publicly obtainable, so no reported association value could be put against a reproduced output. The limitation is on the data-availability side and is no-fault — controlled access is structural, not an authors' defect — so q5/q7 are graded yellow (undetermined) rather than red (fabrication-suspect). The link-mined code (flashpca, only a PCA sub-component) and data accession (GEO GSE107585, a kidney snRNA-seq false positive) are mismatched artifacts, not the real pipeline. No reproduction was attempted and none could be, so the central claim remains untested rather than refuted.

🤝
Reproduced automatically — and fairly

Automated reproduction checks whether a published result can be regenerated from the paper’s described methods and shared data. When something does not reproduce, that is not a claim of error or misconduct — most often it reflects under-described methods, software or environment differences, or gaps in data access, and some of the pre-print papers in the queue may carry issues their authors had no part in. The goal is shared awareness that rigorous, fully-described methods help everyone — never a judgement of any author.

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Reproduction footprint

<synthetic>

Measured resources invested to assess this paper — sanitised (machine class only, no job ids/paths). Compute = HPC accounting (SLURM); tokens = the AI agent's session.

12.7 k
tokens (I/O) · 356.8 k incl. cache
13 min
runtime
Per-job HPC accounting not captured for this run — the runtime shown is the reproduction’s measured wall-clock time.