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Identification of key genes in chickpea transcriptomics and the development of ChickpeaOmicsR as a comprehensive resource to advance breeding and genomic studie

Front Bioinform · 2026
L1 56/100 3/4
⚑ Flagged for review — a reproduced result did not match the reported value

Provisional — an automated or curator check raised a specific concern and points reviewers here. This is NOT a final assessment and not a determination about the authors.

Why this verdict

The main results reproduced, with only marginal, non-material deviations.

Reproduced on the brainbox compute brainarbeit.com
Scoring basis — itemised

Every item that counted toward this verdict, and the exact part of the reproduction that produced it.

Supporting (toward a concern)
Content-critical question only partially held
+2 pts
From: Q5 · Derivability / plausibility 🟡
Content-critical question only partially held
+2 pts
From: Q7 · Core claim 🟡
Content-critical question only partially held
+2 pts
From: Q8 · Severity of the miss (overall human judgment) 🟡
Minor / cosmetic deviation
+1 pts
From: Q6 · Severity of the deviation 🟡
Minor / cosmetic deviation
+1 pts
From: Q2 · Endpoint comparability 🟡
Concordant (toward reproduced)
Code + data deposited & functional
-2 pts
From: Data & code availability Available & functional
Total score +6
✓ What held up
  • Nothing in this column.
What did not (or only partly)
  • 🟡Could not use the authors’ exact input data
  • 🟡Reported values were only indirectly comparable
  • 🔴A deviation arose in the data or preprocessing
  • 🔴A deviation was attributed to the published material
  • 🟡Reported values were not (fully) derivable from the shared data
  • 🟡The deviation was non-trivial in magnitude
  • 🟡The central claim did not (fully) hold under reproduction
  • 🟡Overall, the reproduction showed a material discrepancy
How its reproducibility compares
56/100
Reproducibility score
1.0 SD below mean
vs. all fields · 1173 studies
🎯 Scores higher than 16% of all assessed papers rank 979 of 1173 scored

A 0–100 reproducibility-quality score from the per-question grades, shown as a z-score: standard deviations above (+) or below (−) the mean of comparable assessments.

Reproduction agent’s raw note

Described well enough to verify the resource, NOT to reproduce every reported number 1:1. ChickpeaOmicsR is a resource package that ships the paper's processed pipeline outputs (8 .RData); I re-derived the reported numbers from those shipped objects on «our HPC» rather than re-running HISAT2/HTSeq/DESeq2 on ~205 SRA samples (the package ships the counts, so a raw re-run would not be a 1:1 check and is the deliberate hard-20% skip). RESULT = PARTIAL with notable flags: (1) the 3 named key genes (Fig 5C) are genuine and reproduce 1:1 including exact protein annotations; annotation gene count (24,681) is exact and GWAS gene count (1,050 vs 1,052) within tolerance; the Fig 5C 'five stress conditions' claim holds at FDR<0.05 (2/3 genes in 5 conditions). (2) BUT the headline count-matrix dimension quoted in the README, R/data.R and the paper (28,891 genes x 197 samples) does NOT match the actual shipped CaExpressionCounts (24,702 x 200) -- and no shipped object anywhere contains 28,891 genes; sample count is stated three different ways (197/200/205). (3) The Fig 3A intersection counts (2,910 shared) do not reproduce (got 4,348). These discrepancies are flagged as data-version-drift / possible-fabrication for human audit; the genuine named genes argue against wholesale fabrication. NOT attempted: raw SRA->counts pipeline, PPI 500-node subnetwork selection (unspecified), GWAS GEMMA reanalysis (raw genotypes not shipped), GO enrichment recompute (precomputed table shipped).

These records describe the outcome of reproduction attempts carried out autonomously by brainbox using large language models (LLMs). They are not peer review, not an audit, and not a determination of error or misconduct by any author. A verdict reflects what one attempt could or could not reproduce — which may depend on data access, undocumented parameters, the computing environment, or the depth of effort — and not a judgement of the people who did the work. We can be wrong, and we correct mistakes quickly: every record carries a “report an error” button.

Assessment versions

Every reproduction run is kept as an immutable version — anchored to the data as it stood, with a tamper-evident chain hash. A rerun (e.g. after an author updates a deposit) adds a new version; the previous one stays on record.

  1. v1 current initial assessment Score 56
    assessed: 2026-06-14 ⛓ ceb2803156da
✎ I am an author of this paper

Updated or fixed a deposit, or is there an erratum? Ask us to re-run the metrics. We verify by email first; the new result is published as a new version with full history — nothing is overwritten.

Reason for the rerun

We email you a confirmation link first. The rerun is an objective re-measurement — it cannot change the verdict in your favour, only ask us to look again.

Provenance — full disclosure

When this reproduction was carried out, which methodology version was used, and by whom — so the record can be audited and checked independently.

Reproduced
2026-06-14
Rubric version
v1.0
Assessed by
🤖 AI curator · claude (ai-curator room) · v1.0 · run #1 2026-06-15
no human curator yet
Last updated
2026-08-05

Provisional, curator- or AI-assessed, and independently checkable. A reproduction outcome states what one attempt could reproduce — not a judgement of the authors.

Deep full-text extraction

Model: opus
Founding hypothesis

The study aims to identify genetic mechanisms underlying chickpea (Cicer arietinum L.) responses to diverse biotic and abiotic stressors via transcriptomic profiling, and to develop ChickpeaOmicsR, an integrated R package resource for multi-omics chickpea analysis.

Core claims
  • ChickpeaOmicsR is the first comprehensive/specialized R package integrating transcriptomic, genomic, and proteomic (RNA-seq, GWAS, PPI) data within a unified, reproducible framework and standardizing fragmented chickpea gene nomenclature. resource
  • Each of the six stress/developmental conditions (drought, heat, cold, salinity, Fusarium, developmental stages) triggers distinct molecular pathways. finding
  • Drought and heat stress affected cell wall organization and defense responses. finding
  • Cold stress influenced circadian rhythm genes. finding
  • Fusarium stress involved pathways related to innate immunity and secondary metabolism. finding
  • Developmental stages showed the highest transcriptome variability among the conditions tested. finding
  • An RNA-seq pipeline (fastp/FastQC QC, HISAT2 alignment, HTSeq counts, DESeq2 DEGs, STRING GO/KEGG/PPI) was used to identify stress-responsive genes. method
  • GWAS integration links genetic variants to agronomic traits to support candidate gene identification in chickpea breeding. method
Experimental setups
Assay System Perturbation Readout Platform
bulk RNA-seq (differential gene expression) Cicer arietinum (chickpea), multiple tissues under heat, cold, salinity, drought, Fusarium wilt, and developmental stages abiotic/biotic stress vs. control (heat, cold, salinity, drought, Fusarium) and developmental stage comparison gene-level read counts and differentially expressed genes (FDR<0.01) NCBI SRA datasets; fastp, FastQC, HISAT2, HTSeq, DESeq2 v1.38.3
Gene Ontology / KEGG functional enrichment chickpea candidate genes (Arabidopsis thaliana as annotation model) none enriched GO terms and KEGG pathways STRING database
protein-protein interaction (PPI) network analysis chickpea proteins (Arabidopsis thaliana annotation model) none protein interaction network (nodes/edges) STRING database; Cytoscape
GWAS (genome-wide association study) 679 ICARDA chickpea accessions across 11 environments over 2 years none (natural genetic variation; agronomic traits e.g. DTF, PH, DTM, PPP, HSW, YPP) marker-trait associations (p<0.001) vcf2gwas with GEMMA (MLM Q+K); PLINK PCA; TASSEL filtering
Key results
  • Drought and heat stress affected cell wall organization and defense response pathways
  • Cold stress influenced circadian rhythm genes
  • Fusarium stress involved innate immunity and secondary metabolism pathways
  • Developmental stages showed the highest transcriptome variability among tested conditions
  • Genotyping of 679 accessions yielded ~4 million SNPs, filtered to 92,821 SNPs (MAF>0.05, missing<0.1) 92,821 SNPs from ~4 million
  • Integrated expression dataset compiled covering 28,891 genes across 197 samples (CaExpressionCounts) 28,891 genes; 197 samples
Key statistics
  • count 92,821 SNPs (SNPs retained after TASSEL filtering (MAF>0.05, missing<0.1) from 679 chickpea accessions)
  • count ~4 million SNPs (SNPs from NGS genotyping before filtering)
  • count 28,891 genes across 197 samples (CaExpressionCounts gene expression matrix in ChickpeaOmicsR)
  • pvalue FDR <0.01 (significance threshold for DEGs (Benjamini-Hochberg))
  • pvalue p < 0.001 (significance threshold for GWAS marker-trait associations)
  • count 15 experiments / 6 stress conditions (RNA-seq experiments analyzed across drought, heat, cold, salinity, Fusarium, developmental stages)
  • other 530.8 Mb genome assembly, 120.0x coverage (chickpea genome assembly from NCBI)
  • count 52 samples (28 control vs 24 heat) (heat stress dataset PRJNA748749)

Statistical methods review

Model: sonnet

A neutral, descriptive read of the statistical approach — what was done, and (for shared learning, not as criticism) what could also have been done.

The study performed RNA-seq-based differential gene expression analysis on 15 publicly available chickpea datasets spanning six stress conditions and developmental stages, using DESeq2 (v1.38.3) with Benjamini-Hochberg FDR correction (threshold FDR <0.01). Gene ontology, KEGG pathway enrichment, and protein-protein interaction analyses were conducted via the STRING database using Arabidopsis thaliana as an annotation model. A separate genome-wide association study was conducted on 679 accessions genotyped at 92,821 SNPs using a mixed linear model (Q+K model in GEMMA via vcf2gwas), with population structure (first two PCs) and a kinship matrix included as covariates. Results were reported primarily as lists of significant DEGs and putative trait-associated SNPs, with variance-stabilized expression values visualized in heatmaps.

Replicationbiological Sample sizeBiological replicates per condition listed in Table 1; range from 2 replicates per group (cold, Fusarium, smallest drought datasets) to 28 controls vs. 24 stressed (heat stress); GWAS used 679 accessions across 11 environments over 2 years GroupsStress vs. control for each of six stress types; multiple developmental stage comparisons; 679 accessions for 11 agronomic traits in GWAS Pairingunpaired Randomization/blindingnot stated Dispersionnone Effect sizesyes Confidence intervalsno Multiplicity correctionBenjamini-Hochberg FDR for DEG analysis; no genome-wide multiple-testing correction explicitly applied to GWAS (nominal threshold p <0.001 used)
Statistical tests used
Test Applied to n Assumptions
DESeq2 Wald test with Benjamini-Hochberg FDR adjustment Differential gene expression between stress and control conditions across 15 RNA-seq datasets (six stress types and developmental stages) Varies by dataset: 4–52 samples per experiment as listed in Table 1; analyses restricted to genes with mean count >10 across all samples stated
Mixed linear model (Q+K model, GEMMA via vcf2gwas) Genome-wide association analysis for 11 agronomic traits 679 chickpea accessions, 92,821 SNPs after filtering (MAF >0.05, missing rate <0.1) stated
Functional enrichment analysis (GO and KEGG) via STRING database Gene ontology and pathway analysis of candidate DEG sets per stress condition not stated
Approaches that could also have been used
  • Each of the 15 RNA-seq datasets was analyzed independently with DESeq2, yielding a separate DEG list per dataset even when multiple datasets belonged to the same stress condition
    Could also: Datasets from the same stress condition could be jointly analyzed after batch-effect correction (e.g., ComBat-seq or limma's removeBatchEffect), or combined via a formal meta-analysis framework (e.g., MetaDE, RankProd, or Fisher's combined p-value method) — Joint or meta-analytic approaches pool replication across studies, increasing statistical power for low-replicate datasets (n=2 per group in several experiments) and yielding a single consensus gene list per condition that quantifies cross-study concordance
  • GWAS significance was assessed with a nominal threshold of p <0.001 without a genome-wide multiple-testing correction across the 92,821 SNPs tested
    Could also: A Bonferroni-corrected genome-wide threshold (0.05/92,821 ≈ 5.4×10⁻⁷) or a permutation-based FDR threshold could also be applied — Genome-wide corrections explicitly account for the number of simultaneous hypothesis tests; the authors' relaxed threshold is described as an intentional choice for downstream integrative use, and noting the standard alternative helps readers calibrate confidence in individual locus-trait associations
  • Protein-protein interactions were inferred using the STRING database with Arabidopsis thaliana as the annotation model organism
    Could also: A legume-specific PPI resource (e.g., LegumeIP, SoyBase) or an ortholog-transfer approach mapping chickpea proteins to a more closely related species could also be used — Using a more phylogenetically proximate species or a chickpea-native interaction network reduces the potential for false interactions introduced by evolutionary divergence between Arabidopsis and Cicer arietinum
  • Population structure in the GWAS model was controlled by including only the first two principal components as fixed-effect covariates
    Could also: A larger number of PCs (selected by, e.g., Tracy-Widom test or a scree-plot elbow criterion) or a model-based Q matrix (e.g., from STRUCTURE or ADMIXTURE) could also be incorporated — The optimal number of PCs depends on the complexity of population stratification; in diverse genebank collections such as the ICARDA panel, more than two PCs may be needed to fully account for fine-scale structure and further reduce confounding
  • Heatmap clustering used Pearson correlation distance with complete linkage on rlog-transformed counts
    Could also: Spearman rank correlation distance or Euclidean distance on z-scored values could also be used for hierarchical clustering — Spearman correlation is less sensitive to outlier genes with very high variance; Euclidean distance directly reflects expression-level magnitude differences, which can be informative when the scale of fold change is itself of biological interest
  • The DEG significance threshold was set at FDR <0.01 across all datasets and stress conditions
    Could also: The more common FDR <0.05 threshold used in many transcriptomics studies, or an additional fold-change filter (e.g., |log2FC| ≥1), could also be applied — FDR <0.05 is the default in DESeq2 documentation and widely used as a reference standard; combining an FDR threshold with a minimum fold-change cutoff is also common practice to focus on genes with both statistical significance and a meaningful expression difference
Software: DESeq2 (R) 1.38.3 · R · fastp · HISAT2 · HTSeq · FastQC · pheatmap (R) · STRING database · Cytoscape · vcf2gwas · GEMMA · PLINK · TASSEL

Citation network

Where this publication sits in the reproducibility-weighted citation graph — what it is built on, and what is built on it. Citation data from OpenAlex.

Citations
0
Impact: low
Foundation confidence
None of its references are in our reproducibility record yet — its foundation cannot be assessed.
Topics

No assessed neighbours yet — the network grows as more papers are assessed.

What was reproduced

The exact results taken into scope, with each reported value next to the value our attempt produced.

Scope — pmid-41909810 (ChickpeaOmicsR)

Paper: Identification of key genes in chickpea transcriptomics and the development of ChickpeaOmicsR as a comprehensive resource. Front Bioinform 2026. DOI 10.3389/fbinf.2026.1727493. PMID 41909810.

Code: https://github.com/AlsammanAlsamman/ChickpeaOmicsR (commit 733ff71f4f5ee99f58c3649d9eceeb0bf40b3e61, pushed 2024-12-04). An R resource package that ships the processed pipeline outputs as 8 .RData objects (expression counts, DE significance, annotation, PPI network, GWAS, etc.).

Data: 15 RNA-seq experiments from SRA (PRJNA748749 is the Heat dataset; the package integrates 14 PRJNA studies). Raw reads NOT in repo; the derived count matrix and DE tables are shipped.

Pipeline behind each reported result

  • Raw reads → fastp/fastq-quality-filter → HISAT2 (ref ASM33114v1) → HTSeq counts → DESeq2 v1.38.3 DE (FDR<0.01) → igraph PPI subnetwork + GO/GWAS join.

In scope (reproduced by re-deriving from the shipped objects — "third-party

/own tool on the paper's own data", per brief rule P16/§2)

  • Dimensions of the shipped resource vs the numbers stated in the paper/README (CaExpressionCounts, metadata, annotation, GWAS gene counts). [C1–C4]
  • The 3 named key/hub genes (Fig 5C) — presence + protein-annotation identity. [C5]
  • Hub-gene cross-condition frequency (Fig 5C "five stress conditions"). [C6]
  • Per-condition DEG counts (Table 3) recomputed from CaExpressionSignificance. [C7]
  • DEG intersection across conditions (Fig 3A UpSet counts). [C8]
  • Full PPI network size (context for the Fig 5C subnetwork). [C9]

OUT of scope / not attempted (the hard ~20%, brief §3)

  • Full SRA→counts pipeline (HISAT2/HTSeq/DESeq2 on ~205 samples). The package ships the counts; re-running would not be a 1:1 check of a reported number and is the expensive last 20%. Skipped deliberately.
  • PPI subnetwork selection (Fig 5C "500 nodes / 383 edges"): the rule for reducing the 20k-node network to 500 nodes is not specified → not reproducible.
  • GWAS reanalysis (92,821 SNPs, GEMMA, 679 accessions): raw genotypes not in the repo; only the derived CaGWAS table is shipped.
  • GO enrichment recompute: shipped CaProteinEnrichment is the precomputed table.

All compute ran on «our HPC» (SLURM «job»); RData stayed on «infra», only small JSON results pulled to «host».

Figures / tables: TableFig 5CFig 3A
C1_counts_dims
Reported
28,891 genes x 197 samples
Reproduced
24,702 genes x 200 samples
did not match
C2_metadata
Reported
197 samples / 14 studies / 6 stress conditions
Reproduced
200 samples / 14 PRJNA datasets / 6 conditions (exact set)
partial
C3_annotation_genes
Reported
24,681 genes
Reproduced
24,681 rows
exact
C4_gwas_genes
Reported
1,052 genes
Reproduced
1,050 unique genes
within tolerance
C5_hub_gene_identity
Reported
LOC101503501=Monothiol glutaredoxin-S9; LOC101490854=SRG1; LOC101495985=Thaumatin-like protein 1b (Fig 5C)
Reproduced
all 3 present in shipped Counts/Significance/Annotation; annotations match exactly
exact
C6_hub_five_conditions
Reported
3 key genes frequent under 5 stress conditions (Fig 5C)
Reproduced
FDR<0.05: 2 of 3 DE in exactly 5 conditions, 3rd in 4
partial
C7_DEG_counts_table3
Reported
Heat 658; Cold ~2.2-2.4k; Salinity 20-40; Fusarium 169-802; Dev 54-7150 (Table 3)
Reproduced
Heat 1088; Cold 23/4298/3934; Salinity 5/14; Fusarium 75/823; Dev 50/7411/7774 (FDR<0.01)
partial
C8_intersection_fig3a
Reported
2,910 shared; 2380/484/43/3 across 2/3/4/5 conditions (Fig 3A)
Reproduced
4,348 shared; 3689/590/67/2 across 2/3/4/5 (FDR<0.01)
did not match
C9_ppi_subnetwork
Reported
500 nodes / 383 edges (Fig 5C subnetwork)
Reproduced
full network 20,436 nodes / 746,441 edges; subnetwork rule unspecified
partial

Assessments & scoring basis

Each contributor’s verdict, the per-question basis, and the auditable, itemised worksheet behind it.

🤖 AI curator · claude (ai-curator room) · v1.0 L1 56/100

An automated assessment. It can flag an open question for review but can never, on its own, record a discrepancy verdict (C5) against a paper.

🟡1. Data identity
🟡2. Endpoint comparability
🔴3. Location of the main deviation
🔴4. Cause of the deviation
🟡5. Derivability / plausibility
🟡6. Severity of the deviation
🟡7. Core claim
🟡8. Severity of the miss (overall human judgment)
Scoring basis — itemised

Every item that counted toward this verdict, and the exact part of the reproduction that produced it.

Supporting (toward a concern)
Content-critical question only partially held
+2 pts
From: Q5 · Derivability / plausibility 🟡
Content-critical question only partially held
+2 pts
From: Q7 · Core claim 🟡
Content-critical question only partially held
+2 pts
From: Q8 · Severity of the miss (overall human judgment) 🟡
Minor / cosmetic deviation
+1 pts
From: Q6 · Severity of the deviation 🟡
Minor / cosmetic deviation
+1 pts
From: Q2 · Endpoint comparability 🟡
Concordant (toward reproduced)
Code + data deposited & functional
-2 pts
From: Data & code availability Available & functional
Total score +6

This is a resource paper whose deposited data is genuine: the three named hub genes (LOC101503501/LOC101490854/LOC101495985) reproduce 1:1 with exact protein annotations, the annotation count (24,681) is exact, and GWAS genes (1,050 vs 1,052) are within tolerance — so the central biological conclusion broadly holds and this is not wholesale fabrication. However, the headline count matrix dimension (28,891 genes × 197 samples) stated in three places matches no shipped object (actual 24,702 × 200), and Fig 3A intersection counts (2,910 vs reproduced 4,348) do not reproduce. The deviations sit on the authors'/package side (data-version drift + under-specified DEG thresholds + a value not derivable from any deposit), not our method, and are moderate in severity (structure/order-of-magnitude preserved). Net: partial, flagged for human audit of the unexplained 28,891 figure.

🤝
Reproduced automatically — and fairly

Automated reproduction checks whether a published result can be regenerated from the paper’s described methods and shared data. When something does not reproduce, that is not a claim of error or misconduct — most often it reflects under-described methods, software or environment differences, or gaps in data access, and some of the pre-print papers in the queue may carry issues their authors had no part in. The goal is shared awareness that rigorous, fully-described methods help everyone — never a judgement of any author.

Are you an author? We would genuinely like to hear from you — to clarify the record, add data or code, re-run the pipeline after an accession update, and publish your response right next to the assessment. Everything here is open and auditable.

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Reproduction footprint

claude-opus-4-8

Measured resources invested to assess this paper — sanitised (machine class only, no job ids/paths). Compute = HPC accounting (SLURM); tokens = the AI agent's session.

107.9 k
tokens (I/O) · 6.8 M incl. cache
12 min
runtime · 0.01 CPU-h
2.5 GB
peak RAM
3
HPC jobs
hummel
machine