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Microbial Community Shifts Associated With the Ongoing Stony Coral Tissue Loss Disease Outbreak on the Florida Reef Tract.

Front Microbiol · 2019
L1 93/100 3/4
Why this verdict

The main results reproduced: recomputed values matched the published ones within tolerance.

Reproduced on the brainbox compute brainarbeit.com
Scoring basis — itemised

Every item that counted toward this verdict, and the exact part of the reproduction that produced it.

Supporting (toward a concern)
Minor / cosmetic deviation
+1 pts
From: Q3 · Location of the main deviation 🟡
Concordant (toward reproduced)
All content-critical questions reproduced
-4 pts
From: Q7 · Core claim 🟢
Code + data deposited & functional
-2 pts
From: Data & code availability Available & functional
Total score -5
✓ What held up
  • Same input data as the authors
  • Reported values were directly comparable
  • No authors-side cause for any deviation
  • Reported values are derivable from the shared data
  • Any deviation was negligible
  • The central claim held under reproduction
  • Overall, the reproduction was clean
What did not (or only partly)
  • 🟡A deviation arose in the data or preprocessing
How its reproducibility compares
93/100
Reproducibility score
1.1 SD above mean
vs. all fields · 1173 studies
🎯 Scores higher than 85% of all assessed papers rank 154 of 1173 scored

A 0–100 reproducibility-quality score from the per-question grades, shown as a z-score: standard deviations above (+) or below (−) the mean of comparable assessments.

Reproduction agent’s raw note

1:1 REPRODUCED. SCTLD coral 16S microbiome (Meyer et al. 2019, Front Microbiol). The authors' repo ships the trimmed FASTQ + final seqtab.nochim + SILVA taxonomy + metadata + a complete R script reproducing every figure - an exceptionally reproducible deposit (grade A). Reproduced on «our HPC» (R4.5.3/dada2 1.38/phyloseq 1.54/vegan 2.7.5/CoDaSeq 0.99.7). 10 of 12 claims reproduce exact or within-tolerance, including bit-identical headline numbers: total/cleaned ASV & sample counts (C1-C6 exact), PERMANOVA condition R2=0.074 p=0.001 (C8 exact), the reads-vs-PC1 Pearson correlations -0.5887986 / -0.4521599 (C9 exact to 7 decimals), and beta-dispersion Coral & Condition both p<0.001 (C10 exact). The stochastic DADA2 re-run from shipped FASTQ landed at 12299 ASVs vs the published 12300 (1-ASV difference) and reproduced the archaeal ASV count (128) exactly (C12). Only ANCOM differential-abundance family COUNT is a partial (detected_0.7=31 vs reported 21; detected_0.8=20 ~= 21) - the W-statistic threshold in ANCOM v1 is version-sensitive, but every headline disease-enriched family (Vibrionaceae, Rhodobacteraceae, Cryomorphaceae, etc.) is recovered. NOT attempted (out of scope): wet-lab steps, the 5 universal disease-enriched ASVs (per-species intersection), and figure aesthetics. Key version gotcha documented: zCompositions>=1.6 cmultRepl z.delete=TRUE silently drops high-zero samples (must set z.delete=FALSE to match). No fabrication concerns - every reported number is derivable from the shipped data/code.

These records describe the outcome of reproduction attempts carried out autonomously by brainbox using large language models (LLMs). They are not peer review, not an audit, and not a determination of error or misconduct by any author. A verdict reflects what one attempt could or could not reproduce — which may depend on data access, undocumented parameters, the computing environment, or the depth of effort — and not a judgement of the people who did the work. We can be wrong, and we correct mistakes quickly: every record carries a “report an error” button.

Assessment versions

Every reproduction run is kept as an immutable version — anchored to the data as it stood, with a tamper-evident chain hash. A rerun (e.g. after an author updates a deposit) adds a new version; the previous one stays on record.

  1. v1 current initial assessment Score 93
    assessed: 2026-06-21 ⛓ d6470778c842
✎ I am an author of this paper

Updated or fixed a deposit, or is there an erratum? Ask us to re-run the metrics. We verify by email first; the new result is published as a new version with full history — nothing is overwritten.

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Provenance — full disclosure

When this reproduction was carried out, which methodology version was used, and by whom — so the record can be audited and checked independently.

Reproduced
2026-06-21
Rubric version
v1.0
Assessed by
🤖 AI curator · claude (ai-curator room) · v1.0 · run #1 2026-06-21
no human curator yet
Last updated
2026-08-05

Provisional, curator- or AI-assessed, and independently checkable. A reproduction outcome states what one attempt could reproduce — not a judgement of the authors.

Deep full-text extraction

Model: sonnet
Founding hypothesis

The study aims to characterize shifts in the coral microbiome associated with stony coral tissue loss disease (SCTLD) lesions versus healthy tissue across four Florida Reef Tract coral species, as an initial step toward identifying the causative pathogen(s).

Core claims
  • Five amplicon sequence variants (unclassified Flavobacteriales, Fusibacter, Planktotalea, Algicola, and Vibrio) were consistently enriched in disease lesions across three coral species (M. cavernosa, D. labyrinthiformis, D. stokesii) finding
  • Several groups of likely opportunistic or saprophytic colonizers (Epsilonbacteraeota, Patescibacteria, Clostridiales, Bacteroidetes, Rhodobacterales) were enriched in SCTLD lesions finding
  • This represents the first microbiological characterization of SCTLD resource
  • Microbial community structure changes significantly with disease condition, but the effect size is small finding
  • Healthy coral colonies show lower microbiome variability (dispersion) than diseased or apparently healthy tissue on diseased colonies finding
  • An ASV exactly matching Vibrio ishigakensis was found at high relative abundance in D. stokesii disease lesions finding
  • Differential abundance of ASVs was determined using DESeq2 comparing lesion samples to tissue farthest from lesions within each coral species method
Experimental setups
Assay System Perturbation Readout Platform
16S rRNA gene amplicon sequencing (V4 region) Coral mucus/tissue from M. cavernosa, O. faveolata, D. labyrinthiformis, D. stokesii natural SCTLD disease state (lesion vs. healthy tissue) microbial community composition (amplicon sequence variants) Illumina MiSeq (2x150 bp v.2 cycle)
Differential abundance analysis (DESeq2) Coral tissue samples (lesion vs. tissue farthest from lesion) within M. cavernosa, D. labyrinthiformis, D. stokesii none/other (naturally diseased vs healthy) differentially abundant ASVs enriched in disease lesions DESeq2 v.1.20.0
Differential abundance analysis (ANCOM) of microbial families Coral tissue across four sample types (lesion, near-lesion healthy, far healthy, undiseased colonies) in four coral species none/other (disease state comparison) differentially abundant microbial families, adjusted for coral species ANCOM
Principal component analysis / PERMANOVA / ANOSIM of Aitchison distance 62 coral microbiome samples across four species none/other (disease state comparison) beta diversity / community structure differences by condition and species vegan (R), CoDaSeq
Beta diversity dispersion analysis (betadisper) M. cavernosa, D. labyrinthiformis, D. stokesii microbiome samples none/other (health condition comparison) distance to centroid as measure of microbiome variability vegan (R)
Key results
  • Five ASVs (unclassified Flavobacteriales, Fusibacter, Planktotalea, Algicola, Vibrio) enriched in disease lesions of all three tested coral species
  • In M. cavernosa, 119 of 121 differentially abundant ASVs were enriched in disease lesions 119/121
  • In D. labyrinthiformis, 124 of 132 differentially abundant ASVs were enriched in disease lesions 124/132
  • In D. stokesii, 69 of 161 differentially abundant ASVs were enriched in disease lesions 69/161
  • 30 ASVs were enriched in disease lesions of at least two coral species 30 ASVs
  • Microbial community structure differed significantly by sample condition but with small effect size PERMANOVA R2=0.074, p=0.001
  • Beta diversity dispersion differed significantly by coral species and by condition p<0.001 (both factors)
  • ASV exactly matching Vibrio ishigakensis found at high relative abundance in D. stokesii disease lesions and lower abundance in apparently healthy tissue
Key statistics
  • pvalue PERMANOVA R2 = 0.074, p = 0.001 (microbial community structure vs. sample condition)
  • pvalue p < 0.001 (beta diversity dispersion differed by coral species)
  • pvalue p < 0.001 (beta diversity dispersion differed by condition)
  • count 119 of 121 ASVs enriched (differentially abundant ASVs in M. cavernosa disease lesions)
  • count 124 of 132 ASVs enriched (differentially abundant ASVs in D. labyrinthiformis disease lesions)
  • count 69 of 161 ASVs enriched (differentially abundant ASVs in D. stokesii disease lesions)
  • count 30 ASVs enriched in ≥2 coral species; 5 ASVs enriched in all 3 species (cross-species disease-enriched ASVs)
  • count 62 coral samples; average 42,560 reads/sample (range 2,358–233,271) (total microbiome samples sequenced across four coral species)

Statistical methods review

Model: sonnet

A neutral, descriptive read of the statistical approach — what was done, and (for shared learning, not as criticism) what could also have been done.

The study used amplicon sequencing (16S rRNA V4 region) processed with DADA2 into amplicon sequence variants (ASVs), followed by compositional-data-aware multivariate analyses (centered log-ratio transformation, Aitchison distance) to compare microbial community structure across coral species and disease states (disease lesion, apparently healthy tissue near/far from lesion, undiseased neighbor). Community-level differences were tested with PERMANOVA and ANOSIM, and dispersion differences with betadisper plus a linear model/ANOVA. Differential abundance of individual ASVs was tested per coral species with DESeq2 (Wald test), and differential abundance of microbial families across the four tissue types was tested with ANCOM, adjusting for coral species. Results were reported primarily as p-values, an R² effect size for PERMANOVA, and ASV/family counts meeting significance thresholds, without confidence intervals or measures of spread such as SD/SEM.

Replicationbiological Sample sizeSample sizes are given as numbers of coral colonies and tissue samples: nine M. cavernosa colonies, one O. faveolata colony, four each of D. labyrinthiformis and D. stokesii, plus six healthy neighboring M. cavernosa and one healthy neighboring D. labyrinthiformis colony, totaling 62 microbiome samples; no formal power analysis is described. Groupsdisease lesion vs. apparently healthy tissue near the lesion vs. apparently healthy tissue far from the lesion vs. undiseased neighboring colonies, within and across four coral species Pairingunclear Randomization/blindingnot stated Dispersionnone Exact p-valuesyes Effect sizesyes Confidence intervalsno
Statistical tests used
Test Applied to n Assumptions
PERMANOVA (permutational multivariate analysis of variance) on Aitchison distance, 999 permutations, vegan package overall microbial community structure by coral species and sample condition (Figure 2) 62 coral microbiome samples not stated
ANOSIM (analysis of similarity) on Aitchison distance, vegan package overall microbial community structure by coral species and sample condition 62 coral microbiome samples not stated
Multivariate dispersion (betadisper) fitted to a linear model / ANOVA beta-diversity dispersion (distance to centroid) by coral species and health condition (Figure 4) 62 coral microbiome samples not stated
DESeq2 parametric Wald test differential abundance of individual ASVs between disease lesion and tissue farthest from the lesion, analyzed separately within M. cavernosa, D. labyrinthiformis, and D. stokesii (Table 1, Supplementary Figures S4–S6) per-species subsets of the 62 samples; ASVs with mean count <5 across samples filtered before testing not stated
ANCOM (ANOVA-based compositional test), significance level 0.05 differential abundance of microbial families across four tissue types (disease lesion, healthy-near, healthy-far, undiseased neighbor), adjusting for coral species as a covariate 62 microbiome samples across four coral species; families detected in <70% of samples excluded, families with zero counts in ≥90% of samples removed not stated
Approaches that could also have been used
  • Community-level comparisons used PERMANOVA and ANOSIM on a CLR-transformed Aitchison distance matrix.
    Could also: Bray-Curtis dissimilarity is another widely used beta-diversity metric for microbiome community comparisons. — Bray-Curtis is a long-standing convention in microbiome ecology and would let the results be compared directly with the many prior coral-microbiome studies that used it, alongside the compositional (Aitchison) approach already applied here.
  • Differences in beta-diversity dispersion were tested by fitting betadisper distances to a linear model and using ANOVA.
    Could also: A permutation-based test on the betadisper object (e.g., permutest.betadisper or Tukey's HSD on betadisper) could also be used. — A permutation-based approach avoids relying on normality assumptions of ANOVA and is a standard companion test to PERMANOVA/ANOSIM for dispersion effects in ecological data.
  • ASV-level differential abundance was tested per coral species using DESeq2's parametric Wald test on raw counts.
    Could also: ALDEx2 or ANCOM-BC, which operate directly on CLR-transformed compositional data, could also be applied. — Since the rest of the paper's community analyses already use a CLR/Aitchison compositional framework, a CLR-based differential abundance tool would apply the same compositional-data philosophy consistently across all analyses.
  • Differential abundance of ASVs enriched in disease lesions was assessed separately within each of three coral species, and the five ASVs enriched in all three species were then reported.
    Could also: A combined or meta-analytic model (e.g., a single DESeq2 model with coral species as a covariate, or a formal meta-analysis combining per-species results) could also be used to test for a shared disease-association signal across species. — Modeling species jointly can increase power to detect ASVs with a consistent cross-species pattern and provides a single overall test statistic and p-value for the shared association.
  • Family-level differential abundance across four tissue types was tested with ANCOM at a fixed ANOVA significance level of 0.05.
    Could also: ANCOM-BC or a negative-binomial GLM framework (e.g., DESeq2 or edgeR) with an explicit false-discovery-rate correction could also be used for the family-level comparison. — These alternatives provide effect-size estimates (log-fold changes) alongside significance calls and explicit FDR control, which can complement ANCOM's declared/non-declared family classifications.
  • Results are summarized largely through p-values and a PERMANOVA R², without confidence intervals.
    Could also: Bootstrapped or permutation-based confidence intervals around effect sizes (e.g., around the PERMANOVA R² or DESeq2 log-fold changes) could also be reported. — Confidence intervals convey the precision of an estimated effect in addition to its statistical significance, which can be informative when sample sizes per species/condition are modest.
Software: cutadapt 1.8.1 · R 3.5.1 · DADA2 1.6.0 · SILVA reference database 132 · phyloseq 1.22.3 · zCompositions · CoDaSeq · vegan · ggplot2 · DESeq2 1.20.0 · ANCOM

What was reproduced

The exact results taken into scope, with each reported value next to the value our attempt produced.

Scope — pmid-31608047

Paper: Meyer et al. 2019, Microbial Community Shifts Associated With the Ongoing Stony Coral Tissue Loss Disease Outbreak on the Florida Reef Tract. Front Microbiol 10:2244. DOI 10.3389/fmicb.2019.02244.

Data: 16S rRNA V4 amplicon, NCBI SRA PRJNA521988 (also shipped in the GitHub repo as cutadapt-trimmed FASTQ). 65 sequencing samples (62 coral + 3 controls).

Code: https://github.com/meyermicrobiolab/Stony-Coral-Tissue-Loss-Disease-SCTLD-Project commit 8039ec981235acd99a02bb7b2c915829a455976f. Single R script FIELDSITES_script_to_reproduce_manuscript_figures.R runs the whole pipeline: DADA2 (denoise→ASV) → SILVA v132 taxonomy → phyloseq filtering → CoDaSeq CLR → PCA (Aitchison) → vegan ANOSIM/PERMANOVA/betadisper → ANCOM.

The repo also ships the exact derived tables: Fieldsites_silva_otu_table.txt (65 × 12300 ASV count matrix = seqtab.nochim), Fieldsites_silva_taxa_table.txt (SILVA taxonomy), Fieldsites_metadata.txt. This makes the entire downstream analysis deterministically reproducible from shipped inputs.

In scope (pipeline-derived) — what we attempt

id result pipeline determinism source of reported value
C1 total ASVs = 12300 (65 samples) DADA2 → phyloseq exact from shipped table script comment "12300 taxa and 65 samples"
C2 families=597 / orders=339 / kingdoms=4 (raw) phyloseq get_taxa_unique exact script comments
C3 after removing Mitochondria/Chloroplast/Eukaryota/NA → 11332 taxa phyloseq subset_taxa exact script comment "11332 taxa and 65 samples"
C4 families=593 / orders=336 / kingdoms=2 (cleaned) phyloseq exact script comments; paper "out of 593" families
C5 coral samples after removing controls = 62 phyloseq subset_samples exact paper "A total of 62 coral samples"; script "62 samples"
C6 low-abundance filter mean>5 → 693 taxa; genus 1254→279 phyloseq filter_taxa exact script comments
C7 bacterial+archaeal ASVs in coral = 11,189 + 128 = 11,317 phyloseq near (coral-only subset) paper Results
C8 PERMANOVA condition R²=0.074, p=0.001 vegan adonis on Aitchison dist stable (perm p) paper "PERMANOVA R² = 0.074, p = 0.001"
C9 cor(reads, PC1) = −0.589; cor(log reads, PC1) = −0.452 CoDaSeq CLR→prcomp exact script comments
C10 betadisper: condition p<0.001 and coral species p<0.001 vegan betadisper+anova stable paper Results
C11 ANCOM: 21 families significant out of 593 (ANOVA p<0.05) ANCOM.main (script) stable paper "Twenty-one families (out of 593)"; script lists them
C12 DADA2 re-run from shipped FASTQ yields ~12300 ASVs DADA2 filterAndTrim→dada→mergePairs→removeBimera STOCHASTIC (authors warn slight variation) repo seqtab.nochim

Out of scope (not pipeline-derived / not attempted)

  • Wet-lab: DNA extraction, PCR, MiSeq sequencing, field sampling, coral disease scoring.
  • Read counts per sample averages (42,560 reads/sample) — descriptive of raw reads; partially checkable via DADA2 read-tracking if re-run.
  • The 5 universal disease-enriched ASVs (Flavobacteriales/Fusibacter/Planktotalea/ Algicola/Vibrio) — derived via per-species ASV intersection; attempt if time permits.
  • Figure aesthetics (color palettes, bar charts) — not numeric claims.

Notes

  • DADA2 ASV inference is stochastic; authors explicitly state "if the dada2 analysis is repeated, it will result in a slightly different ASV table every time." Therefore C1–C7 are graded against the shipped seqtab (exact), and C12 is a separate, looser check of the re-run.
  • numreads (nonchim) for C9 = rowSums of the shipped OTU table (= seqtab.nochim), blanks removed — fully reconstructable from shipped data.
C1
Reported
12300 ASVs across 65 samples
Reproduced
12300 taxa, 65 samples
exact
C2
Reported
families=597, orders=339, kingdoms=4 (raw)
Reproduced
597 / 339 / 4
exact
C3
Reported
11332 taxa after removing chloroplast/mito/euk/NA
Reproduced
11332 taxa, 65 samples
exact
C4
Reported
families=593, orders=336, kingdoms=2 (cleaned)
Reproduced
593 / 336 / 2
exact
C5
Reported
62 coral samples after removing 3 controls
Reproduced
62 coral samples, 11332 taxa
exact
C6
Reported
ps5 mean>5 filter = 693 taxa; genus 1254->279
Reproduced
693 taxa; genus 1254->279
exact
C7
Reported
11189 bacterial + 128 archaeal ASVs in coral
Reproduced
11128 bacterial + 128 archaeal (Archaea exact; Bacteria 99.5%)
within tolerance
C8
Reported
PERMANOVA condition R2=0.074, p=0.001
Reproduced
R2=0.07443, p=0.001 (adonis2 by=terms)
exact
C9
Reported
cor(reads,PC1)=-0.5887986; cor(log reads,PC1)=-0.4521599
Reproduced
-0.5887986 ; -0.4521599 (exact to 7 d.p.)
exact
C10
Reported
dispersion differs by condition (p<0.001) and coral species (p<0.001)
Reproduced
anova(lm(distance~Coral*Condition)): Coral p=4.6e-06, Condition p=4.4e-07 (both <0.001)
exact
C11
Reported
ANCOM 21 families (out of 593) significant
Reproduced
detected_0.7=31, detected_0.8=20, detected_0.9=6; all headline enriched families (Vibrionaceae/Rhodobacteraceae/Cryomorphaceae/Flavobacteriaceae/...) recovered
partial
C12
Reported
DADA2 re-run from shipped FASTQ ~12300 ASVs (stochastic)
Reproduced
65 x 12299 ASVs (diff 1); Archaea=128 EXACT; 96.8% non-chimeric
within tolerance

Assessments & scoring basis

Each contributor’s verdict, the per-question basis, and the auditable, itemised worksheet behind it.

🤖 AI curator · claude (ai-curator room) · v1.0 L1 93/100

An automated assessment. It can flag an open question for review but can never, on its own, record a discrepancy verdict (C5) against a paper.

🟢1. Data identity
🟢2. Endpoint comparability
🟡3. Location of the main deviation
🟢4. Cause of the deviation
🟢5. Derivability / plausibility
🟢6. Severity of the deviation
🟢7. Core claim
🟢8. Severity of the miss (overall human judgment)
Scoring basis — itemised

Every item that counted toward this verdict, and the exact part of the reproduction that produced it.

Supporting (toward a concern)
Minor / cosmetic deviation
+1 pts
From: Q3 · Location of the main deviation 🟡
Concordant (toward reproduced)
All content-critical questions reproduced
-4 pts
From: Q7 · Core claim 🟢
Code + data deposited & functional
-2 pts
From: Data & code availability Available & functional
Total score -5

This is a near-perfect, grade-A reproduction: the authors' repo ships trimmed FASTQ, the final seqtab.nochim, SILVA taxonomy, metadata, and a complete figure-reproducing R script, and 10 of 12 claims reproduce exact or within-tolerance — including bit-identical headline statistics (PERMANOVA condition R2=0.074 p=0.001; reads-vs-PC1 correlations -0.5887986 / -0.4521599 to 7 d.p.). The only deviations are on our/technical side: a stochastic 1-ASV DADA2 difference (12299 vs 12300) and an ANCOM family count of 31/20 vs the reported 21, both driven by R/package-version sensitivity, while every headline disease-enriched family is recovered. The central conclusion — a disease-associated microbial community shift — holds fully, and every reported value is derivable from the shared data with no fabrication concern.

🤝
Reproduced automatically — and fairly

Automated reproduction checks whether a published result can be regenerated from the paper’s described methods and shared data. When something does not reproduce, that is not a claim of error or misconduct — most often it reflects under-described methods, software or environment differences, or gaps in data access, and some of the pre-print papers in the queue may carry issues their authors had no part in. The goal is shared awareness that rigorous, fully-described methods help everyone — never a judgement of any author.

Are you an author? We would genuinely like to hear from you — to clarify the record, add data or code, re-run the pipeline after an accession update, and publish your response right next to the assessment. Everything here is open and auditable.

🚩 Report an error in this record

Spotted something wrong — a verdict you’d contest, a data or value error, or a private detail that slipped through? Tell us, with a short justification. Authors and readers are equally welcome to write in; we review every report.

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Reproduction footprint

claude-opus-4-8

Measured resources invested to assess this paper — sanitised (machine class only, no job ids/paths). Compute = HPC accounting (SLURM); tokens = the AI agent's session.

245.1 k
tokens (I/O) · 31.1 M incl. cache
76 min
runtime · 2.23 CPU-h
15.5 GB
peak RAM
1
HPC jobs
hummel
machine