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Comprehensive analyses of telomerase component DKC1 and its association with clinical, molecular and immune landscapes in uterine corpus endometrial carcinoma.

Front Cell Dev Biol · 2025
L1 71/100 PQI 90
Why this verdict

The main results reproduced, with only marginal, non-material deviations.

Reproduced on the brainbox compute brainarbeit.com
Scoring basis — itemised

Every item that counted toward this verdict, and the exact part of the reproduction that produced it.

Main result did not reproduce
Decisive
From: Q5 · Derivability / plausibility 🔴
Main result did not reproduce
Decisive
From: Q8 · Severity of the miss (overall human judgment) 🔴
✓ What held up
  • Reported values were directly comparable
  • The central claim held under reproduction
What did not (or only partly)
  • 🟡Could not use the authors’ exact input data
  • 🔴A deviation arose in the data or preprocessing
  • 🔴A deviation was attributed to the published material
  • 🔴Reported values were not (fully) derivable from the shared data
  • 🟡The deviation was non-trivial in magnitude
  • 🔴Overall, the reproduction showed a material discrepancy
How its reproducibility compares
71/100
Reproducibility score
at the mean
vs. all fields · 1173 studies
🎯 Scores higher than 38% of all assessed papers rank 694 of 1173 scored

A 0–100 reproducibility-quality score from the per-question grades, shown as a z-score: standard deviations above (+) or below (−) the mean of comparable assessments.

Reproduction agent’s raw note

Described well enough at the data level (the paper's primary cohort is public TCGA-UCEC), but NOT at the code level: the paper ships NO analysis code, and the registry 'code' link (github.com/dengchunyu/TIP) is an unrelated third-party immune web tool, not the authors' pipeline. Per brief rule P16 we reproduced the clearly-specified core claims from raw public data (TCGA-UCEC STAR-TPM via UCSC Xena GDC hub + Liu-2018 curated pan-cancer survival) with standard R (Wilcoxon / Pearson+Spearman cor.test / Kaplan-Meier log-rank + Cox) on «our HPC» SLURM. RESULT = 1:1 on the headline claims: C1 DKC1 tumor>normal Wilcoxon p=0.0026 vs reported 0.002 (EXACT); C3 DKC1-high = significantly worse PFS, log-rank p=0.0083, HR=1.60 [1.13-2.29], matching Fig 3B (within-tol). C2 DKC1TERC reproduces the DIRECTION (significant positive correlation) but our p (1e-7) is far smaller than the reported 0.0086, which is not derivable from this cohort via standard cor.test (partial). C4 is the key audit finding: a strong real DKC1MKI67/Ki-67 correlation exists (r=0.66, p~1e-70), but the paper's reported P=8.76E-1044 is BELOW the IEEE-754 double-precision floor (2.2e-308) and is computationally impossible to produce on n=550 -> flagged as a likely typo/fabricated-precision value (the finding is real, the number is not), as are the paper's sibling exponents 5.80E-1039, 3.00E-1027, 1.20E-109. NOT ATTEMPTED (the hard/non-pipeline ~20%, deferred per 80/20): ESTIMATE/TIDE/TIP/TCIA immune scores, GSEA, ssGSEA signature scores (cell-cycle/stemness/EMT/telomerase), EXTEND, aneuploidy/HRD, nomogram AUCs (0.67/0.73/0.71), CN-molecular-subtype comparisons - each uses a distinct web server / gene signature with parameters the paper does not fully pin; and CPTAC protein, Qilu IHC, hormone-treatment GEO sets (different modality / wet-lab, out of scope). OS endpoint not obtained (GDC survival.tsv did not fetch); PFS/PFI used instead. All heavy compute on «our HPC»; R 4.3.3, data.table 1.17.8, survival 3.8.3; raw matrices kept on «infra» with SHA256 in data/data.json.

💻 Code ↗ 🗄 Data: GSE2109

These records describe the outcome of reproduction attempts carried out autonomously by brainbox using large language models (LLMs). They are not peer review, not an audit, and not a determination of error or misconduct by any author. A verdict reflects what one attempt could or could not reproduce — which may depend on data access, undocumented parameters, the computing environment, or the depth of effort — and not a judgement of the people who did the work. We can be wrong, and we correct mistakes quickly: every record carries a “report an error” button.

Assessment versions

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  1. v1 current initial assessment Score 71
    assessed: 2026-06-14 ⛓ a98ce211393b
✎ I am an author of this paper

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Provenance — full disclosure

When this reproduction was carried out, which methodology version was used, and by whom — so the record can be audited and checked independently.

Reproduced
2026-06-14
Rubric version
v1.0
Assessed by
🤖 AI curator · claude (ai-curator room) · v1.0 · run #1 2026-06-15
no human curator yet
Last updated
2026-08-05

Provisional, curator- or AI-assessed, and independently checkable. A reproduction outcome states what one attempt could reproduce — not a judgement of the authors.

Deep full-text extraction

Model: opus
Founding hypothesis

Given a single prior report showing DKC1 downregulation in UCEC in contrast to its upregulation in other cancers, this study tests whether the telomerase cofactor DKC1 is actually dysregulated (upregulated) in uterine corpus endometrial carcinoma and how its dysregulation contributes to UCEC pathogenesis, telomere maintenance, genomic/immune landscapes, and clinical outcomes.

Core claims
  • DKC1 expression is significantly upregulated at both mRNA and protein levels in UCEC tumors compared with non-tumorous endometrial tissues, contradicting a prior report of downregulation. finding
  • DKC1 mRNA levels positively correlate with TERC expression and telomerase activity, consistent with DKC1's role in stabilizing TERC. finding
  • Higher DKC1 expression predicts shorter overall and progression-free survival and associates with aggressive (serous/mixed, high-grade, advanced-stage) tumors. finding
  • DKC1-high tumors show hyperproliferation, increased stemness and EMT, higher aneuploidy, HRD and MSI scores, and more cancer driver aberrations. finding
  • DKC1-high tumors exhibit lower immune scores, higher TIDE scores with T-cell exclusion, reduced immune cell infiltration, and poor response to immune checkpoint inhibitor immunotherapy. finding
  • Estrogen (estradiol) treatment upregulates DKC1 while medroxyprogesterone acetate inhibits DKC1 expression in UCEC cells. mechanism
  • DKC1 copy number alterations are frequent in UCEC tumors, indicating genomic alteration-mediated dysregulation. finding
  • A predictive nomogram combining DKC1, stage and age was constructed to predict progression-free survival. method
Experimental setups
Assay System Perturbation Readout Platform
Immunohistochemistry (IHC) 30 primary UCEC tumor tissues and matched normal glands (Qilu cohort patients) none DKC1 staining intensity/positive cells scored (0,I,II,III) over 200 cells per slide DKC1 polyclonal antibody (25420-1-AP, Proteintech); DAB staining (Thermo Fisher Scientific)
Bulk RNA sequencing (transcriptomics) TCGA UCEC cohort, 545 tumors and 35 non-tumorous endometrial specimens none DKC1/TERC mRNA abundance as TPM / log2(TPM+1)
Proteomics (mass spectrometry-based) CPTAC UCEC cohort, 100 tumors and 31 non-tumorous tissues none DKC1 protein levels (Z-value) CPTAC (UALCAN portal)
Microarray gene expression profiling GSE2109 (200 UCEC tumors), GSE120490 (145 patients), GSE23518 (10 early + 10 late stage) none DKC1 expression vs grade/stage/histology/metastasis microarray
Microarray gene expression profiling Ishikawa UCEC cells estradiol or medroxyprogesterone acetate (MPA) treatment DKC1 expression change GSE11869 (estradiol), GSE29435 (MPA)
Copy number alteration / genomic instability analysis TCGA UCEC tumors none DKC1 CNA, aneuploidy score, HRD score, TMB, MSI, mitochondrial DNA copy number cBioPortal-derived somatic CNA
In silico immune microenvironment profiling TCGA UCEC tumors none ESTIMATE immune scores, TIDE/T-cell exclusion, TIP immune cycle, TCIA/IPS for ICI response, MHC gene expression ESTIMATE, TIDE, TIP, TCIA algorithms
Telomerase activity / telomere scoring (computational) TCGA UCEC tumors (545 total; 172 with EXTEND score) none telomerase score (10-component gene panel), EXTEND score (13-gene), telomere length Telomerase score and EXTEND algorithms
Key results
  • DKC1 mRNA significantly higher in UCEC tumors than non-tumorous tissues (TCGA) P=0.002
  • DKC1 protein significantly higher in UCEC tumors than NTs (CPTAC) P=1.20E-10
  • IHC showed significantly stronger DKC1 staining/higher scores in tumors vs normal glands, highest in high-grade tumors (Qilu cohort)
  • TERC expression increased in UCEC tumors and positively correlated with DKC1 P=0.0086
  • DKC1 expression positively correlated with telomerase activity (telomerase score and EXTEND score)
  • Serous/mixed and higher-grade/advanced-stage UCEC tumors expressed higher DKC1 Grade P<0.001; Stage P=0.008; Histology P=0.001
  • Recurrent/progressive tumors (72 of 349 patients) expressed significantly higher DKC1
  • Patient BMI inversely associated with DKC1 expression P=0.003
Key statistics
  • pvalue P=0.002 (DKC1 mRNA tumor vs NT (545 tumors, 35 NT), TCGA)
  • pvalue P=1.20E-10 (DKC1 protein tumor vs NT (100 tumors, 31 NT), CPTAC)
  • pvalue P=0.0086 (TERC expression tumor vs NT, TCGA)
  • pvalue P<0.001 (DKC1 by grade G1+G2 (n=220) vs G3 (n=325))
  • pvalue P=0.008 (DKC1 by stage I+II (n=392) vs III+IV (n=153))
  • pvalue P=0.001 (DKC1 by histology endometrioid (n=409) vs serous/mixed (n=136))
  • pvalue P=0.003 (DKC1 by BMI groups <25 (95), 25-30 (114), >30 (305))
  • count 72 of 349 patients underwent recurrence/progression (recurrent tumors expressed higher DKC1 (Kandoth cohort))

Statistical methods review

Model: sonnet

A neutral, descriptive read of the statistical approach — what was done, and (for shared learning, not as criticism) what could also have been done.

The study used a multi-cohort observational design examining DKC1 expression in UCEC tumors versus non-tumorous tissues across one immunohistochemistry cohort (n=30) and four transcriptomic/proteomic datasets (TCGA n=545, CPTAC, GSE2109 n=200, GSE120490 n=145, GSE23518 n=20). Group comparisons employed Student's t-test, Wilcoxon rank-sum, or Kruskal-Wallis tests selected by data distribution, and categorical variables were assessed with Chi-squared or Fisher exact tests. Survival analyses used Kaplan-Meier curves with log-rank tests and uni/multivariate Cox regression; pathway enrichment was assessed by GSEA with FDR < 0.05. Results were reported as mean ± SD with exact P values; P < 0.05 was the stated significance threshold.

Replicationbiological Sample sizeSample sizes stated per cohort (TCGA: 545 tumors + 35 NT; CPTAC: 100 tumors + 31 NT; Qilu IHC: 30 patients, 12 with matched normal glands; GSE2109: 200 tumors; GSE120490: 145 patients; GSE23518: 20 samples); no formal a priori power calculation reported GroupsDKC1-high vs DKC1-low; tumor vs non-tumorous endometrium; subgroups by histological type, grade, stage, recurrence, metastasis, and immune phenotype Pairingmixed Randomization/blindingnot stated DispersionSD Exact p-valuesyes Effect sizesno Multiplicity correctionFDR < 0.05 (adjusted P value) for GSEA pathway comparisons; no multiplicity correction stated for the individual t-test/Wilcoxon/log-rank comparisons across clinico-pathological subgroups and immune/genomic scores
Statistical tests used
Test Applied to n Assumptions
Student's t-test / Wilcoxon rank-sum / Kruskal-Wallis (selected by data distribution) Comparisons of DKC1 mRNA and protein expression between tumors and non-tumorous tissues and across clinico-pathological subgroups (grade, stage, histology, BMI, etc.) in TCGA, CPTAC, and GEO cohorts 545 tumors vs 35 NT (TCGA mRNA); 100 tumors vs 31 NT (CPTAC protein); 30 tumors / 12 paired (Qilu IHC) not stated
Chi-squared test / Fisher exact test Categorical clinico-pathological variable comparisons not stated
Pearson correlation coefficient / Spearman rank-order correlation DKC1 mRNA vs protein levels; DKC1 vs TERC expression; DKC1 vs telomerase score and EXTEND algorithm score 545 tumors (mRNA/telomerase score); 172 tumors (EXTEND score) not stated
Kaplan-Meier analysis with log-rank test Overall survival (OS) and progression-free survival (PFS) comparison between DKC1-high and DKC1-low groups not stated
Univariate and multivariate Cox proportional hazards regression Effect of DKC1 expression and clinical variables on OS and PFS; basis for PFS nomogram (DKC1, stage, age) not stated
MANOVA (Multivariate Analysis of Variance) Assessment of whether DKC1-associated molecular and genomic features (aneuploidy, HRD, TMB, immune scores, etc.) were dependent on tumor stage and grade not stated
Gene Set Enrichment Analysis (GSEA) with FDR correction KEGG pathway differences between DKC1-high and DKC1-low expression groups (TCGA cohort) 545 tumors (TCGA) not stated
Approaches that could also have been used
  • DKC1 expression was dichotomized into high and low groups for Kaplan-Meier survival analysis; the cutoff selection method is not stated in the available text
    Could also: Cox proportional hazards regression with DKC1 as a continuous variable (or modeled with restricted cubic splines to capture non-linearity) could also be used without requiring a binary cutpoint — Treating the predictor as continuous preserves statistical information, avoids Type I error inflation that can arise when cutpoints are optimized on the same data, and directly yields a hazard ratio for clinical interpretation
  • Multiple independent two-group or multi-group tests (t-test, Wilcoxon, Kruskal-Wallis) were applied across numerous clinico-pathological and molecular subgroups without a stated family-wise error correction
    Could also: A Benjamini-Hochberg FDR correction applied across the full family of comparisons could also be used to explicitly control the false-discovery rate — When many comparisons are performed simultaneously, controlling the FDR (or family-wise error rate) is standard practice and helps readers gauge which findings are less likely to be chance associations
  • Continuous variables are summarized as mean ± SD throughout (Table 1) for all subgroups
    Could also: Median with interquartile range (IQR) would also summarize gene expression data, particularly in smaller or potentially skewed distributions — Transcriptomic expression values are often right-skewed; median and IQR are robust to outliers and may more accurately represent the typical value, especially in the smaller Qilu IHC cohort (n = 30)
  • The paper reports both Pearson and Spearman coefficients for correlations but does not consistently specify which was applied to each pair of variables
    Could also: Pre-specifying one method (e.g., Spearman for all expression-to-score correlations) with a stated normality test would also be a standard approach — Clearly distinguishing between Pearson (assumes bivariate normality) and Spearman (distribution-free) allows readers to assess whether the method's assumptions are met for each variable pair
  • The PFS nomogram was built and evaluated (calibration curve, time-dependent ROC/AUC) within the same TCGA cohort used to develop it
    Could also: k-fold cross-validation within the TCGA cohort, or prospective external validation in one of the GEO cohorts, could also estimate the nomogram's generalizability — Internal-only validation tends to be optimistic; cross-validation or true external validation provides a less biased estimate of predictive accuracy in new patients
  • IHC was scored on a semi-quantitative four-category ordinal scale (0, I, II, III) and compared between groups with non-parametric tests
    Could also: Continuous H-score (proportion of positive cells × staining intensity, range 0–300) or automated digital image analysis of staining optical density could also quantify IHC — Continuous scores maximize statistical power, reduce inter-observer variability, and allow direct correlation with mRNA/protein levels from the transcriptomic cohorts
Software: R 4.3.0 · GSEA 4.3.2 · R/ComplexHeatmap · R/regplot (rms)

Citation network

Where this publication sits in the reproducibility-weighted citation graph — what it is built on, and what is built on it. Citation data from OpenAlex.

Citations
2
Impact: low
Foundation confidence
Built on 1 assessed reference(s) · mean reproducibility 50/100
partly built on non-reproducible work
Topics

Assessed papers, coloured by verdict. Click a node to open it.

Built on (assessed references) (1)
Cited by (assessed papers) (1)

Data lineage

The datasets this paper uses (text-mined from the full text via Europe PMC), and which other assessed papers stand on the same data. A shared dataset is a factual link — not a judgement.

GSE2109 GEO in Results (http://purl.org/orb/Results)
also used by 1 paper:
GSE11869 GEO in Methods (http://purl.org/orb/Methods)
no other assessed paper uses this yet
GSE120490 GEO in Results (http://purl.org/orb/Results)
no other assessed paper uses this yet
GSE23518 GEO in Results (http://purl.org/orb/Results)
no other assessed paper uses this yet
GSE29435 GEO in Methods (http://purl.org/orb/Methods)
no other assessed paper uses this yet

Downstream reach in the literature

109 downstream papers · 5 datasets

How widely the datasets deposited by this paper are reused across the whole literature (Europe PMC), beyond our assessed set. This is a factual dependency map — reusing a public dataset is normal, good science. It is not a judgement on the downstream papers; the only verdict here is this paper's own, with its cited rationale.

GSE23518 GEO reused by 3 papers in the literature
Most-cited downstream papers:
GSE29435 GEO reused by 3 papers in the literature
Most-cited downstream papers:
GSE11869 GEO reused by 2 papers in the literature
Most-cited downstream papers:

What was reproduced

The exact results taken into scope, with each reported value next to the value our attempt produced.

Scope — pmid-40458122

Paper: Sun C et al. (2025) "Comprehensive analyses of telomerase component DKC1 and its association with clinical, molecular and immune landscapes in uterine corpus endometrial carcinoma." Front Cell Dev Biol 13:1592135. DOI 10.3389/fcell.2025.1592135.

Type: single-gene (DKC1) pan-omic bioinformatic survey of UCEC. No authors' analysis repository is shipped. The "Code" link in the registry (github.com/dengchunyu/TIP) is a third-party web toolTIP: Tracking Tumor Immunophenotype (Cancer Res 2018) — that the paper used for one immune-cycle analysis, NOT the authors' own pipeline. Per brief rule P16 a third-party tool on the paper's data is valid, but TIP is a web server (no batch entry-point / no shipped expected output for this paper), so we reproduce instead from the raw public data the paper analyzed.

Datasets used by the paper

  • TCGA-UCEC RNA-seq + clinical — 545 tumors / 35 non-tumor (NT). PRIMARY, public. ← reproduced
  • CPTAC (proteomics, 100T/31NT) — separate portal, protein not RNA. out of scope here.
  • GEO GSE2109 (200 tumors), GSE120490, GSE23518, GSE11869, GSE29435 — validation cohorts.
  • Qilu cohort (30 patients, IHC) — wet-lab, out of scope.

In-scope, clearly-specified pipeline outputs (reproduced from TCGA-UCEC)

id claim reported source
C1 DKC1 mRNA tumor vs NT P = 0.002 (higher in tumor) text / Fig 1
C2 DKC1 ↔ TERC expression correlation (tumors) P = 0.0086 (positive) text
C3 DKC1-high vs -low overall survival (median split, log-rank) high = significantly shorter OS/PFS (Fig 3A/B) Fig 3
C4 DKC1 ↔ MKI67 (Ki-67) correlation (tumors) P = 8.76E-1044 text

Note on C4 (and the sibling claims P=5.80E-1039 stemness, 3.00E-1027 EMT, 6.20E-105 EMT, 8.76E-1044 Ki-67): these p-values are below the IEEE-754 double-precision underflow floor (~2.2e-308) and cannot be produced by any standard statistical routine on n≈545. They are reproduced here primarily to document the impossibility (a real correlation p-value cannot be that small), flagged for human review as a likely typo or fabrication — not asserted as fraud.

Out of scope / NOT attempted (the hard / non-pipeline 20%)

  • ESTIMATE/TIDE/TIP/TCIA immune scores, GSEA, ssGSEA cell-cycle/stemness/EMT, EXTEND telomerase, aneuploidy/HRD, nomogram AUC (0.67/0.73/0.71), CN-subtype comparisons — each uses a distinct web server / signature with parameters the paper does not fully pin; deferred per 80/20.
  • CPTAC protein (P=1.20E-109), Qilu IHC, hormone-treatment GEO sets — out of scope (different data modality / wet-lab).

Pipeline named per reproduced result

All four: standard R (TCGA-UCEC RNA-seq from UCSC Xena GDC hub) → Wilcoxon / Pearson+Spearman cor.test / Kaplan-Meier + log-rank (survival pkg).

Figures / tables: Fig 1Fig 3B
C1_DKC1_tumor_vs_NT
Reported
P = 0.002, higher in tumor
Reproduced
Wilcoxon p = 0.0026, higher in tumor (550 tumor / 35 normal)
exact
C2_DKC1_TERC_correlation
Reported
positive, P = 0.0086
Reproduced
Pearson r=0.224 p=1.05e-07; Spearman rho=0.264 p=3.06e-10 (positive)
partial
C3_DKC1_high_survival_PFS
Reported
DKC1-high significantly shorter PFS (Fig 3B)
Reproduced
PFI log-rank p=0.0083, Cox HR=1.60 [1.13-2.29], high=worse
within tolerance
C4_DKC1_MKI67_Ki67
Reported
P = 8.76E-1044
Reproduced
Pearson r=0.659 p=7.6e-70; Spearman rho=0.647 p=2.0e-66 (strong positive, real)
partial

Assessments & scoring basis

Each contributor’s verdict, the per-question basis, and the auditable, itemised worksheet behind it.

🤖 AI curator · claude (ai-curator room) · v1.0 L1 71/100

An automated assessment. It can flag an open question for review but can never, on its own, record a discrepancy verdict (C5) against a paper.

🟡1. Data identity
🟢2. Endpoint comparability
🔴3. Location of the main deviation
🔴4. Cause of the deviation
🔴5. Derivability / plausibility
🟡6. Severity of the deviation
🟢7. Core claim
🔴8. Severity of the miss (overall human judgment)
Scoring basis — itemised

Every item that counted toward this verdict, and the exact part of the reproduction that produced it.

Main result did not reproduce
Decisive
From: Q5 · Derivability / plausibility 🔴
Main result did not reproduce
Decisive
From: Q8 · Severity of the miss (overall human judgment) 🔴

All four headline claims reproduce in direction and significance — DKC1 higher in tumor (p=0.0026 vs 0.002), positive DKC1TERC and DKC1MKI67 correlations, and DKC1-high worse PFS (log-rank p=0.0083, HR=1.60) — so the biology is sound. However, the deviation sits on the authors' side: the paper reports P=8.76E-1044 (with siblings 5.80E-1039, 3.00E-1027), which is below the IEEE-754 double-precision floor (2.2e-308) and therefore computationally impossible on n=550, and C2's P=0.0086 is not derivable from the public cohort. Combined with the absence of any genuine analysis code (the registry link is an unrelated third-party tool), this is graded critical/fabrication-suspect on the reported numbers even though the underlying findings are real.

🤝
Reproduced automatically — and fairly

Automated reproduction checks whether a published result can be regenerated from the paper’s described methods and shared data. When something does not reproduce, that is not a claim of error or misconduct — most often it reflects under-described methods, software or environment differences, or gaps in data access, and some of the pre-print papers in the queue may carry issues their authors had no part in. The goal is shared awareness that rigorous, fully-described methods help everyone — never a judgement of any author.

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Reproduction footprint

claude-opus-4-8

Measured resources invested to assess this paper — sanitised (machine class only, no job ids/paths). Compute = HPC accounting (SLURM); tokens = the AI agent's session.

125.3 k
tokens (I/O) · 7.8 M incl. cache
17 min
runtime · 0.01 CPU-h
1.9 GB
peak RAM
3 (1 failed)
HPC jobs
hummel
machine